An anime movie based on the boy band Strawberry Prince.

Second-year high school student Yatora Yaguchi is a delinquent with excellent grades, but is unmotivated to find his true calling in life. Yatora spends his days working hard to maintain his academic standing while hanging out with his equally unambitious friends. However, beneath his carefree demeanor, Yatora does not enjoy either activity and wishes he could find something more fulfilling. While mulling over his predicament, Yatora finds himself staring at a vibrant landscape of Shibuya. Unable to express how he feels about the unusually breathtaking sight, he picks up a paintbrush, hoping his thoughts will be conveyed on canvas. After receiving praise for his work, the joy he feels sends him on a journey to enter the extremely competitive Tokyo University of the Arts—a school that only accepts one in every two hundred applicants. Facing talented peers, a lack of understanding of the fine arts, and struggles to obtain his parents’ approval, Yatora is confronted by much adversity. In the hopes of securing one of the five prestigious spots in his program of choice, Yatora must show that his inexperience does not define him.

Finally, the day of Mai's high school graduation has arrived. While Sakuta eagerly waits for his girlfriend, an elementary schooler who looks exactly like her appears before him. Suspicious, and for all the wrong reasons... Meanwhile, Sakuta and Kaede's father suddenly calls, saying that their mother wants to see her daughter. She was hospitalized because Kaede's condition had been too much for her to bear, so what could she possibly want now?

It's the Taisho era—a time filled with the scent of romance, and people's hearts are still filled with compassion. Unzen, Sukayu, Unohana, Oshamanbe, and Aso are all ordinary (?) male students at Mayunan High School, who spend their days lazing around in the clubroom of the "do-nothing club" Earth Defense Club, and live together in the Kurodama dormitory. One day, while the five are taking a morning bath at the Kurodama dormitory, they notice something slimy moving in the bathtub. When they finally manage to catch it, it turns out to be a talking Pallas's cat named Nuru! Nuru has come from the distant future with a serious mission to inform them that the Earth is facing an impending threat of destruction. While he says he wants to protect the Earth, he says he's only come to warn them of the crisis, and that it's up to the five of them to decide whether or not they want to fight for the Earth... Isn't this just an attempt to dodge responsibility? The Defense Club members are suspicious. "If either is fine with you, I'll do it! That's more fashionable!" Unzen enthusiastically says. The five decide to fight with free will, and Nuru transforms them into the Fashionable Romantic Troupe using cat science. Meanwhile, a mysterious trio called the Bankara New Elite Squad are scheming in the shadows. They are plotting to take over the Earth.